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10.1007/s40620-017-0392-z 139 MasseyA.KiffinR.CuervoA

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The findings have been cited by dozens of subsequent studies, including the 2018 Pickart & Margolina review of GHK-Cu regeneration science and the 2025 Kuceki study combining copper peptides with microneedling

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A 2026 systematic review confirmed that L-Citrulline, especially when combined with antioxidants, increases nitric oxide bioavailability and reduces oxidative stress in endothelial cells